Well,
smart does refer to pain as one use- but a quick look at Webster shows that many other uses such as: spirited, bright, witty, mentally alert, shrewd, sharp in scheming, and on and on. Webster then goes to list a doz or so words using "Smart" as a prefex such as smartypants, smartmouth, smarten, smarty, etc- all denoting definitions dealing with the common usage of "bright, witty, etc"
So I would think that if asked, most would define "smart" as something similar to the above.
but as long as I looking at Webster:
LEARNED: 1) to gain knowledge or understanding by study, instruction or EXPERIENCE 2) to come to be able to 3) to come to realize
RESPONSE: 1) something constituting a reply or a reaction 2) the activity or inhibition of previous activity of an organism or any of its parts resulting from stimulation.
Easy enough to put them together and see that a
learned response is, to paraphrase, "The ability to gain knowledge through experience of a previous activity".
and lets look at
"keen senses"amoung the uses of "
keen" I find (not counting wailing at funerals) listed "sharp" "quick responsivness" "extremely sensitive in perception" "clever" "to grasp" "to comprehend"
and "
senses" I find means: "a meaning conveyed" "the faculty of perceiving" "sentient, intellegent" it goes on to say" SENSE implys a reliable ability to judge and decide.."
so "
keen senses" would certainly imply intellegence- by its very definition..
and "
WARY" means "Careful, aware, attentive" "marked by KEEN caution, cunning and watchful prudence"
CUNNING can then be defined as "skillful, of knowelge, learning" "exhibiting skill" "CLEVER"
So round and round we go....word games sure are fun
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It all comes down to those coyotes can drive ya nuts! 
