I have never found a detailed description of how the lugs at the muzzle work, but since they only come into play when the barrel is at full recoil, their purpose must be to retard or stop the barrel from coming off the carriage if the recuperator looses air or oil pressure.
The picture of the 75 with the barrel in full recoil probably had a failed recuperator, maybe that was why it was captured. The barrel should not be in this position if the recuperator is working properly.
There are a number of French 75's around the US and a very nice one along with a limber, is located at the Harry Truman Library and Museum, in Independence. Missouri. Presented to the library by the French Government. Truman commanded a battery of French 75's in WW1.